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Department of Human Genetics



Acting Chairperson: Donald R. Mattison, MD
Department Office: A300 Crabtree Hall
(412) 624-3018, FAX (412) 624-3020
E-mail: JNORBUT@HELIX.HGEN.PITT.EDU

  • Purpose
    The Department of Human Genetics provides graduate training in the fields of human genetics and genetic counseling. The mission of the department is to discover new knowledge about the genetic determinants of human health and disease through basic and applied research; to educate students, trainees, and other interested persons in that knowledge; and to apply that knowledge to improve the health of populations, families, and patients.

    The human genetics curriculum provides an understanding of basic genetic mechanisms affecting human genes, chromosomes, cells, individuals, and populations of organisms in both normal and disease states. Public health applications are achieved through the detection and quantitation of the effects of intrinsic factors such as age and sex, exposure to extrinsic agents such as chemical pollutants or therapeutic agents, and by understanding gene-environmental interactions in determining the distribution of health and disease in the human population. The underlying focus is on the theory and techniques for collecting, analyzing, and interpreting quantitative data.

  • Major Educational Areas and Programs
    Courses offered by the department address the areas of human population and quantitative genetics, genetic toxicology, biochemical and molecular genetics, cytogenetics, and genetic counseling.

    The principal objective of the courses in human genetics is to train students to critically examine the role of genes and genetic variation in determining the distribution of health and disease in the general population. To achieve this objective, training is provided in both experimental and statistical approaches to the direct detection or estimation of the impact of genes on the health of individuals, families, and populations. Such approaches include the evaluation of the relative roles of genetic and environmental factors and their interaction in determining the distribution of disease in the population, so-called "ecogenetics." The program provides for concentrations in two areas: human genetics (MS, PhD) and genetic counseling (MS).

  • Human Genetics
    This area is concerned with the study of the mechanisms of genetic variability and its impact on health at the population level. An important component is the study of the fraction of genetic variability that leads directly to disease or determines an individual's susceptibility to diseases caused by pathogens or adverse environments.

  • Genetic Counseling
    The objectives of the program are: (1) To provide a balanced program of study integrating courses in molecular genetics, medical genetics, and psychosocial and multicultural counseling including biomedical ethics; (2) to provide extensive direct patient contact experience in a variety of clinical placements so that the student gains an appreciation of how the practicing genetic counselor functions in different work settings; and (3) to prepare students at the Master of Science level for entering the profession of genetic counseling and assuming the role of a professional in medical and academic settings.

    This is a full-time, two-year Master of Science program. The program follows the Graduate School of Public Health core requirements supplemented by human genetics courses. The course work occurs in the first year of study followed by a ten-month clinical rotation at nearby hospitals. Students must take all core courses of the School and may take electives outside the course requirements of the genetic counseling program. A comprehensive written examination is administered in November of the second year of training, and an oral exam for counseling skills is administered in January of the second year. All students must complete a thesis for graduation. The program balances studies in genetics, molecular biology, epidemiology, biostatistics, and counseling intervention. Clinical internships involving laboratory experience and direct patient contact are incorporated into the program. The curriculum exceeds the minimum degree requirements of the Graduate School of Public Health and the American Board of Genetic Counseling (ABGC). All graduates are eligible for ABGC certification.

  • Research
    Research in the Department of Human Genetics includes studies of basic genetic mechanisms of segregation and recombination; family and population studies of normal and disease phenotypes; chromosome structure and chromosomal mechanisms in disease; physical and genetic mapping (genomics); and genetic epidemiology with special reference to common diseases and the importance of genes to public health, interaction of genes with the environment, assessment of genetic risk, and the detection of genetic disease. Application of the knowledge is explored with research in ethics, genetic counseling and screening, as well as in therapy by gene transfer. The focus of faculty research is on human genetics but includes experimental studies in appropriate non-human animals and studies on evolution.

  • Faculty Research Interests

    Research interests represented in the department are:
    Primary Faculty

    Dr. Aston: Genetic epidemiology, statistical methods and their application to human population genetics, quantitative genetics and gene mapping, computation and informatics in genetics

    Dr. Bahnson: Gene therapy, gene transfer methods, hematopoiesis, analytical and regulatory quality control

    Dr. Barranger: Molecular biology of lysosomal function and disorders, enzyme structure and translocation/membrane integration signals, gene therapy, transgenic models

    Dr. Deka: Anthropological genetics, population genetics, comparative genetics, gene mapping

    Dr. Ferrell: Biochemical and molecular genetics, study of the role of genes in determining the distribution of common disease in the population, gene mapping

    Ms. Gettig: Clinical genetic counseling, teratogens, professional education, genetic health care delivery

    Dr. Gollin: Cytogenetics; chromosome structure, organization, and function in healthy and diseased individuals; cancer cytogenetics; gene mapping by fluorescence in situ hybridization

    Dr. Kamboh: Molecular and biochemical genetics; evaluation and identification of the role of genes in determining genetic susceptibility/resistance to cardiovascular disease and Alzheimer's disease in the general population using a combination of contemporary biochemical, molecular, and statistical techniques

    Dr. Law: Application of molecular genetics to better understand complex human diseases, genetic alterations associated with human cancer, and cell cycle regulation

    Dr. Li: Human population genetics, segregation analysis, path analysis, experimental designs, paternity problems, DNA profile match problems

    Dr. McPherson: Clinical aspects of genetic disorders, inheritance of renal agenesis/dysplasia, inheritance of malignant hyperthermia, craniofacial dysplasia

    Dr. Mehdi: Molecular biology, in vitro mutagenesis and expansion

    Dr. Mulvihill: Clinical genetics of cancer, disease gene mapping, syndromes of multiple malformations, genetic and reproductive consequences of medical therapeutics

    Dr. Parker: Ethical issues in genetic research, provision of genetic services, and clinical practice

    Dr. Sanghera: Application of molecular genetics and statistical techniques to identify the role of genes in cardiovascular disease in the general population

    Dr. Shekhter-Levin: Cytogenetics, genetic mechanisms in human cancer with emphasis on hematologic malignancies, clinical cytogenetics

    Dr. Weeks: Linkage analysis, statistical methods in human genetics, population genetics, gene mapping, DNA match problems

    Joint Faculty

    Dr. Bansal: Reproductive genetics, fetal therapy, in utero cell transplantation, fetal gene therapy

    Dr. Gill: Immunogenetics, study of the role of genes in determining susceptibility to infectious and autoimmune diseases, reproductive genetics, ecogenetics

    Dr. Glorioso: Molecular biology, vectors for human gene therapy, especially herpes virus

    Dr. Gorin: Clinical and molecular genetics of human retinal diseases, clinical ophthalmic genetics, cloning of genes for animal models of retinal disorders, gene therapy

    Dr. Hoffman: Genetic linkage, pathogenic, and animal studies of neuromuscular diseases

    Dr. Hogge: Gene therapy, genetics of schizophrenia

    Dr. Latimer: DNA repair genes and mechanisms, especially in common cancers

    Dr. Marazita: Birth defects, especially clefting

    Dr. Nimgoankar: Genetics of schizophrenia

    Dr. Perlin: Mapping of chromosomes, molecular diagnostics and mapping of disease genes

    Dr. Richard: Molecular genetics of neuropsychiatric disorders, genetic and physical maps of human chromosomes 11 and 13

    Dr. Surti: Cytogenetics, cytogenetic mechanisms in human germ cells and somatic cells and their relationship to genetic disease, reproductive genetics

    Dr. Suzuki: Genes in dental disorders, especially periodontal disease

    Dr. Trucco: Genetics and immunogenetics of human histocompatibility antigens, markers for susceptibility to autoimmune diseases, MHC restricted and non-restricted cytotoxicity

    Dr. Wald: Diagnosis and treatment of human radiation overexposure, assessment of environmental chromosome damage in peripheral blood and bone marrow cells, automated dicentric scoring

    Dr. Wenger: Cytogenetic and molecular studies of fragile X syndrome and Prader-Willi syndrome, clinical genetics

    Adjunct Faculty

    Ms. Berry: Genetic counseling

    Ms. Clemens: Genetic counseling

    Mr. Coss: Genetic counseling

    Dr. Filkins: Methods of prenatal diagnosis, including chorionic villus sampling; prospective epidemiological study of exposures in the first trimester of pregnancy; fetal therapy including fetal immunization

    Ms. Fraer: Maternal serum alpha-fetoprotein screening

    Dr. Jackson: Genetic counseling

    Mr. Kerr: Genetic counseling

    Ms. LeChein: Genetic counseling

    Ms. Linn: Genetic counseling

    Dr. Naylor: Biochemical genetics, biochemical detection of human disease and genetic screening, biochemical mechanisms in human metabolic disorders

    Dr. Nicholas: Computer analysis of sequence data

    Ms. Nicklaus: Clinical counseling, teratogen counseling

    Ms. Penles-Stadler: Clinical counseling, especially for familial breast cancer

    Mr. Smith: Genetic counseling

    Dr. Weiss: Anthropogenetics, role of genes in determining the distribution of disease in North American aboriginal populations

  • Qualifications for Admission
    Students in human genetics should have completed courses in calculus, biology, biochemistry, and chemistry. In some cases deficiencies can be made up after admission.

    All accepted students are eligible for financial aid, which may include scholarships, graduate student assistantships, and teaching fellowships.

  • Course Offerings

    HUGEN 2017 HUMAN GENETICS: 3 credits; Spring Term; prerequisite: one course in basic genetics; admission by permission of instructor
    Provides an introduction to basic genetic principles and their application to human health and disease.

    HUGEN 2021 SPECIAL STUDIES: Variable credits; every term
    Qualified students may undertake advanced work or research with the approval and under the guidance of a member of the faculty.

    HUGEN 2022 HUMAN POPULATION GENETICS: 3 credits; Spring Term; prerequisites: one course in statistics, one in genetics
    Deals with general principles of population genetics, especially those applicable to human populations.

    HUGEN 2023 POPULATION GENETICS AND EVOLUTION: 3 credits; Fall Term; (alternate years; given in 1997-98); prerequisites: BIOST 2042, an advanced course in genetics
    Discusses various ways of establishing genetic polymorphisms in populations, including mutation, selection, and subdivision of populations. The consequences of non-random mating and finite population size are also covered. Experimental data from natural populations are included to illustrate the predictions from theory.

    HUGEN 2025 HUMAN GENETICS SEMINAR: 1 credit; Fall Term and Spring Term
    Active researchers present current genetics methodology, theory, and findings

    HUGEN 2026 SPECIAL STUDIES: HUMAN GENETICS: 3 credits; every term; admission by permission of instructor
    Designed to provide advanced undergraduates and graduate students with directed, intensive training in laboratory, statistical, or clinical research methods relevant to human genetics. These will be specialized skills not available through regularly taught courses in the University. Each special study is designed in consultation with an individual member of the Human Genetics faculty. Course performance will be judged by the preparation of a final written report to the supervising faculty member.

    HUGEN 2030 CHEMICAL MUTATION OF HUMAN CHROMOSOMES: 3 credits; Spring Term; prerequisites: organic chemistry, biochemistry, basic genetics, human genetics, biostatistics
    A structure-reactivity approach is taken regarding the interaction of potentially mutagenic/carcinogenic chemicals with human chromatin. Methods for assessing the genetic toxicity of environmental, industrial, and chemotherapeutic agents are discussed. Emphasis is on utilization of in vitro human and other mammalian cytogenetic assays and in vivo animal models for extrapolation to human populations.

    HUGEN 2031 CHROMOSOMES AND HUMAN DISEASES: 3 credits; Fall Term; prerequisite: HUGEN 2017 or equivalent.
    Course familiarizes graduate students and fellows in Human Genetics, Genetic Counseling, or other biomedical areas with the field of cytogenetics, the study of chromosomes, including the relationship of chromosome abnormalities to human disease. Topics covered include, cell structure and function, the cell cycle, chromosome structure and segregation, the mitochondrial genome, classical and molecular cytogenetic methods, nomenclature, sex chromosomes and sex determination, sex chromosome abnormalities, aneuploidy/numerical chromosome abnormalities, structural aberrations, clinical implications of chromosome abnormalities, mosaicism, high-resolution chromosome studies, contiguous gene syndromes, uniparental disomy, genomic imprinting, the Human Genome Project, mapping, cancer cytogenetics, and ethical issues in cytogenetics.

    HUGEN 2032 CYTOGENETIC TECHNIQUES: 2 credits; Spring Term
    Familiarizes graduate students in Human Genetics, Genetic Counseling, or other biomedical areas with genetic techniques including cell culture, cytogenetic harvesting, slide making, staining, chromosome banding, chromosome analysis, automated karyotyping, karyotype interpretation and reporting, and molecular genetic techniques, including Southern blotting, PCR sequencing, as applied to problems in human molecular genetics, including diagnosis, family studies, linkage analysis, and trinucleotide repeats.

    HUGEN 2033 QUANTITATIVE GENETICS: 3 credits; Fall Term; (alternate years; given in 1996-97); prerequisites: HUGEN 2022; BIOST 2043; course in computer science; permission of instructor
    An advanced course for those who have a major interest in quantitative aspects of genetics, namely, in the genetic epidemiology of human diseases.

    HUGEN 2034 INTRODUCTION TO HUMAN BIOCHEMICAL AND MOLECULAR GENETICS: 3 credits; Fall Term; prerequisites: HUGEN 2017 or equivalent; biochemistry
    An introductory course in human biochemical and molecular genetics with a focus on common diseases (hypertension, coronary artery disease, etc.) that require the interaction of inherited disease susceptibility and environmental exposure that results in a clinical phenotype. This course addresses the question of the biochemical and molecular basis of disease susceptibility in the general population.

    HUGEN 2035 PRINCIPLES OF GENETIC COUNSELING: 3 credits; Fall Term; prerequisites: HUGEN 2017, BIOST 2041; or permission of instructor
    Provides basic didactic training in genetic counseling with particular reference to its applications in public health programs and clinical settings.

    HUGEN 2036 GENETIC COUNSELING INTERNSHIP: 4 credits/term; Fall Term and Spring Term; prerequisites: HUGEN 2017, 2022, 2031, 2035; BIOST 2014, 2041
    Provides practical application of the fundamentals of genetic counseling. Involves both the observation of and participation in at least 50 counseling sessions at various clinics. Literature review and searches pertinent to each patient will be necessary for adequate preparation. Weekly case presentations and/or lectures will also be included.

    HUGEN 2038 INTERVENTION SKILLS FOR GENETIC COUNSELORS: 3 credits; Spring Term; prerequisite: registration for HUGEN 2035
    A 22-week seminar course focusing on the understanding of theories of intervention, skill development, and application to genetic counseling. The course aims at sensitizing students to the ethical dilemmas faced by affected families and health care providers.

    HUGEN 2039 RISK CALCULATION IN GENETIC COUNSELING: 1 credit; Summer Session II; prerequisites: HUGEN 2022; BIOST 2041
    Provides training in calculating risk of diseases or carrier status in a variety of genetic counseling situations by learning to identify the sources of risk in the counselee's personal and family history and to analyze and synthesize a single overall risk of disease from the competing risks.

    HUGEN 2040 MOLECULAR BASIS OF HUMAN INHERITED DISEASE: 3 credits; Spring Term; prerequisites: undergraduate-level biology and biochemistry
    Provides an up-to-date overview of the most common and biologically informative human inherited disorders and integrates clinical descriptions with recent genetic, molecular genetic, and biochemical insights. Disorders covered include lysosomal storage disorders, neuromuscular diseases, organic acidopathies, amino acidopathies, neurofibromatosis, cystic fibrosis, neurodegenerative, and ophthalmic disorders. Current techniques of gene mapping, cloning, transfer, and expression will be integrated into the overviews.

    HUGEN 2041 ADVANCED MEDICAL ETHICS: 3 credits; Spring Term; cross-listed as HPS 2664
    An advanced treatment of significant problems in medical ethics. Topics may include euthanasia, rights to health care, competency, allocation of resources, and other issues of medical ethics. Students are expected to have completed at least one introductory course in medical ethics as well as one general ethics course prior to enrollment.

    HUGEN 2042 IMMUNOGENETICS: 3 credits; Spring Term; prerequisites: previous course in genetics and a knowledge of biochemistry
    Deals in depth with the use of immunological and molecular biological techniques to study genetics, mainly, of cell antigens. The emphasis will be on the use of semiological and biochemical tools to identify and isolate cell surface antigens and on their functional significance. The genetics and chemistry of the immunoglobulins will also be covered in detail. The molecular biology underlying the control of cell surface antigens and immunoglobulins will be discussed.

    HUGEN 2043 TOPICS IN MEDICAL ETHICS: 3 credits; Spring Term
    Provides a forum to consider special topics in clinical, medical, and research ethics in a small group.

    HUGEN 2047 CLINICAL GENETICS CASE CONFERENCE: 1 credit; Spring Term, Fall Term, and Summer Sessions; prerequisite: in the genetic counseling program or permission of instructor
    With clinical cases and specimens from various clinical genetics service units, this seminar illustrates and provides insights into the biologic, medical, ethical, and emotional aspects of genetic disorders.

    HUGEN 3010 RESEARCH AND DISSERTATION FOR DOCTORAL DEGREE: Variable credits; hours to be arranged

    FTDR 0000 FULL-TIME DISSERTATION STUDY: Every term
    Doctoral candidates who have completed all credit requirements for the degree, including any minimum dissertation requirements, and are working full-time on their dissertations may register for this course. While the course carries no credits and no grade, students who enroll in "Full-time Dissertation Study" are considered by the University to have full-time registration status.

  • Faculty

    Acting Chairperson
    DONALD R. MATTISON, Dean, Professor of Environmental and Occupational Health and Obstetrics and Gynecology (School of Medicine) and Acting of department. MD, College of Physicians and Surgeons, Columbia University

    Emeritus Professor
    CHING CHUN LI, Emeritus University Professor of Biometry and Human Genetics. PhD, Cornell University

  • Faculty with Primary Appointments

    Professors

    JOHN A. BARRANGER, Professor of Human Genetics; Professor of Pediatrics and Professor of Molecular Genetics and Biochemistry (School of Medicine). MD, PhD, University of Southern California

    ROBERT E. FERRELL, Professor of Human Genetics. PhD, University of Texas

    MOHAMMAD ILYAS KAMBOH, Associate Professor of Human Genetics (promotion pending). MS, University of the Punjab; PhD, The Australian National University

    JOHN J. MULVIHILL, Professor of Human Genetics; Professor of Pediatrics and Professor of Molecular Genetics and Biochemistry (School of Medicine). MD, University of Washington School of Medicine


    Associate Professors

    RANJAN DEKA, Assistant Professor of Human Genetics. MS, PhD, Dibrugarh University

    SUSANNE M. GOLLIN, Associate Professor of Human Genetics. MS, PhD, Northwestern University

    ELIZABETH MCPHERSON, Associate Professor of Human Genetics. MS, University of Wisconsin; MD, University of Washington, Seattle

    DANIEL E. WEEKS, Assistant Professor of Human Genetics (promotion pending). MS, PhD, University of California, Los Angeles

    Assistant Professors

    CHRISTOPHER ASTON, Assistant Professor of Human Genetics. PhD, Australian National University

    ALFRED BAHNSON, Research Assistant Professor of Human Genetics. MS, PhD, University of Pittsburgh

    ELIZABETH GETTIG, Assistant Professor of Human Genetics. MS, Sarah Lawrence College

    JOHN C. LAW, Assistant Professor of Human Genetics. MS, PhD, University of Pittsburgh

    HAIDER MEHDI, Assistant Professor of Human Genetics. MS, PhD, Aligargh Muslim University

    LISA PARKER, Assistant Professor of Human Genetics. MA, PhD, University of Pittsburgh

    DHARAMBIR SANGHERA, Research Associate. PhD, Guru Nanak Dev University

    SOFIA SHEKHTER-LEVIN, Assistant Professor of Human Genetics. MD, ScD, PhD, Pavlov Institute of Physiology of the Academy of Sciences, USSR

    Research Associates

    CHANDRAMOHAN S. ISHWAD, Research Associate. MS, PhD, Bombay University

    FU SHENG WEI, Research Associate. PhD, Beijing University

    JING-FANG WEI, Research Associate. MS, PhD, Beijing University; Chinese Academy of Medical Science

    ZI CHEN ZHANG, Research Associate. MS, MD, Wuhan Medical College

  • Faculty with Secondary Appointments

    Professors

    THOMAS J. GILL III, Professor of Pathology (School of Medicine); Professor of Human Genetics. MA, MD, Harvard University

    JOSEPH C. GLORIOSO, Professor of Molecular Genetics and Biochemistry (School of Medicine); Professor of Human Genetics. PhD, Louisiana State University

    JON B. SUZUKI, Dean of the School of Dental Medicine and Professor of Dental Medicine; Professor of Human Genetics. DDS, Loyola University; PhD, Illinois Institute of Technology

    MASSIMO TRUCCO, Hillman Professor of Pediatric Immunology (School of Medicine); Professor of Human Genetics. MD, University of Torino

    NIEL WALD, Professor of Environmental and Occupational Health and Human Genetics; Professor of Radiology (School of Medicine). MD, New York University

    Associate Professors

    ERIC HOFFMAN, Associate Professor of Molecular Genetics and Biochemistry (School of Medicine); Associate Professor of Human Genetics. PhD, Johns Hopkins University

    WILLIAM A. HOGGE, Associate Professor of Obstetrics, Gynecology, and Reproductive Sciences (School of Medicine); Associate Professor of Human Genetics. MD, University of Virginia

    MARY L. MARAZITA, Associate Professor of Dental Medicine (School of Dental Medicine); Associate Professor of Human Genetics. PhD, University of North Carolina

    URVASHI SURTI, Associate Professor of Pathology (School of Medicine); Associate Professor of Human Genetics. MS, Carnegie Mellon University; PhD, University of Pittsburgh

    SHARON WENGER, Associate Professor of Pediatrics (School of Medicine); Associate Professor of Human Genetics. MS(Hyg), PhD, University of Pittsburgh

    Assistant Professors

    VIJAYA BANSAL, Assistant Professor of Obstetrics, Gynecology, and
    Reproductive Sciences (School of Medicine); Assistant Professor of Human Genetics. MS, MD, University of Delhi; MS, University of Pittsburgh

    MICHAEL B. GORIN, Assistant Professor of Ophthalmology (School of Medicine); Assistant Professor of Human Genetics. MD, PhD, University of Pennsylvania

    JEAN LATIMER, Assistant Professor of Obstetrics, Gynecology, and Reproductive Sciences (School of Medicine). PhD, State University of New York at Buffalo

    VISH NIMGOANKAR, Assistant Professor of Psychiatry (School of Medicine); Assistant Professor of Human Genetics. MD, Christian Medical College

    MARK W. PERLIN, Assistant Professor of Human Genetics. PhD, MD, PhD, Carnegie Mellon University

    CHARLES W. RICHARD III, Assistant Professor of Psychiatry (School of Medicine); Assistant Professor of Human Genetics. MD, PhD, Ohio State University

  • Faculty with Adjunct and Other Appointments

    Professor
    KENNETH WEISS, Adjunct Professor of Human Genetics. Professor of Anthropology, Pennsylvania State University. PhD, University of Michigan (Ann Arbor)

    Associate Professors


    KAREN FILKINS, Clinical Associate Professor of Human Genetics. MD, University of Medicine and Dentistry of New Jersey

    EDWIN A. NAYLOR, Associate Professor of Human Genetics. PhD, Utah State University; MPH, University of Pittsburgh

    HUGH B. NICHOLAS, JR. Associate Professor of Human Genetics. PhD, University of Oklahoma


    Assistant Professor

    CHRISTANN JACKSON, Clinical Assistant Professor of Human Genetics. MD, University of Pittsburgh

    Instructors

    DEBRA BERRY, Department of Reproductive Genetics, West Penn Hospital and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh

    MICHELE CLEMENS, Department of Genetics, Magee-Womens Hospital and Adjunct I Instructor of Human Genetics. MS, University of Pittsburgh

    LUANNE M. FRAER, Department of Genetics, Magee-Womens Hospital and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh

    MICHAEL KERR, March of Dimes and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh

    KAY LECHIEN, Department of Reproductive Genetics, West Penn Hospital and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh

    KATHY LINN, Department of Reproductive Genetics, West Penn Hospital and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh

    EDWARD SMITH, Department of Genetics, Magee-Womens Hospital and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh

    MONA PENLES-STADLER, Department of Genetics, Magee-Womens Hospital and Adjunct Instructor of Human Genetics. MS, University of Pittsburgh


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